Abstract
Antonio García-Bellido was a pioneer of developmental biology in Spain who had the rare ability to see beyond the trees and grasp not just the forest, but entire landscapes. Intellectually and personally, he was indefatigable, uncompromising, and wholly committed to science. He led his trainees by example in his unflagging demands on honoring the data above all else. Antonio discovered the existence of “developmental compartments” in the wing of Drosophila and proposed that these territories corresponded to the realm of action of homeotic genes. He was a fierce defender of hypotheses that he thought best explained available data but did not cling blindly to ideas when they no longer served to understand the phenomena at hand. With his distinctive style he founded a successful school of Drosophila developmental geneticists that will sorely miss his unparalleled passion for science.
Acknowledgments
Author contributions
J.F.d.C., L.A.B.-L., A.B., and C.E. wrote the paper.
Competing interests
The authors declare no competing interest.
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Copyright © 2026 the Author(s). Published by PNAS. This article is distributed under Creative Commons Attribution-NonCommercial-NoDerivatives License 4.0 (CC BY-NC-ND).
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Published online: June 24, 2026
Published in issue: June 30, 2026
Acknowledgments
Author contributions
J.F.d.C., L.A.B.-L., A.B., and C.E. wrote the paper.
Competing interests
The authors declare no competing interest.
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Antonio García-Bellido: A brief history of flies, lineages, and a new developmental genetic logic, Proc. Natl. Acad. Sci. U.S.A.
123 (26) e2614026123,
https://doi.org/10.1073/pnas.2614026123
(2026).
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